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Twigs,
adapted from
Woody Plants of the Blue Ridge
by Ron Lance. Used by permission.
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see photographic examples, click one of the cameras in the list of
botanical terms. |
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The current
or most recent woody growth, forming the ends of a stem, is the
twig.
The previous season's growth (last year's twig) is the branchlet,
sometimes called a 2nd-year stem.
The branch is 3 years or more in age.
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A spur
shoot is knobby, short, and very slow-growing. It usually produces
flowers or clustered leaves and lies along the side of a branchlet
or branch. Sometimes it may be spine-tipped, or merely have a bud
at the tip.
A culm
is the vertical woody stem of bamboos, which have no twigs. The
word cane is also used, but cane is
also applied to the stems of the genus Rubus, which have
true twigs and branchlets.
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A node
is the portion of a twig where vascular tissue exits and enters;
this is where leaves arise.
Internodes
are intervals of twig between nodes.
Where a leaf
falls from the twig, a leaf scar
is left behind. Inside the leaf scar the vascular tissue pattern
is seen, usually as dots or bundles of dots. These are bundle
scars, aiding species identification during Winter.
Buds
are also found at nodes, usually in axil of leaf petiole and twig
or hidden under petiole base, or sunken in twig.
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Position
of buds: Where the twig ends growth by forming a bud on the
very tip, a terminal bud is present.
When the growing tip withers or falls away, one of the side buds
(or lateral buds) ends up being
close to the twig tip and substitutes as a terminal bud for next
year's growth. These are pseudo-terminal
or false-terminal buds;
look for the telltale branch scar
or branch stub alongside it, opposite from leaf scar. This is where
the twig tip has fallen.
Lateral buds
can be superposed (one above
the other) or collateral (side
by side) on the same node. Often, one of these accessory buds is
for flower production and may look different than other lateral
vegetative buds.
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Buds can be
sunken or submerged
in the twig (not visible), naked or scaled.
A naked
bud has the first set of leaves folded and visible as the outside
of the bud, or a heavy layering of hair obscures any distinction
of scales.
Scaled
buds have one or more bud scales (which are modified
leaves) to protect embryonic leaves Inside. When 2-scaled, the scales
may be valvate, or meet like two
cupped hands. Imbricate scales
overlap at edges.
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Other twig
features:
Superficially,
twigs may be lined (raised ridges
extending down from leaf scars) or angled
(not rounded in cross-section). Zig-zag
twigs are bent in alternating angles from one node to the next.
Stipules
are leaf-like appendages found in some plants, usually at base of
petiole or twig nodes. They leave various-shaped scars, or harden
into spines.
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Armed
twigs have sharp thorns, spines, or prickles
Thorns
are outgrowths of the wood of the twig.
Spines are usually associated with
nodes and are modified stipules or twig tips.
Prickles are epidermal outgrowths,
such as in roses and blackberries.
Bristles
are trichomes ("hairs") from the epidermis; stiff but
not hardened and sharp.
Lenticels
are dotlike pores for gas exchange. They can be tiny or conspicuous,
raised and corky, or elongated horizontally.
Aerial
rootlets are thickened fibers for climbing, found on some
vines.
Tendrils
are twining petiole, stipule, or twig modifications that aid
climbing in some vines.
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| Pith
is the soft central core or interior of twigs. Its shape is seen in
cross-section, its texture by splitting or halving the twig lengthwise
using a sharp knife. |
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